Hongye Su, Tao Chang, Li Ma, Zhonghua Zhang, Ruimin Qin, Haze Adi, Jingjing Wei, Shan Li, Xue Hu, Zhengchen Shi, Huakun Zhou (2026). Long-term nitrogen addition of different types and levels altered the relationship between species diversity and ecosystem multifunctionality in alpine grasslands. Soil Ecology Letters, Vol. 80, No. 20.DOI
Jie-xia Li, Qin-Xue Wang, Xi-Lai Li, Zifeng Ma, Yingnian Li, Huakun Zhou (2026). Sensitive response range of alpine meadow degradation on the Qinghai–Tibetan Plateau: Nonlinear fragmentation under pika and mowing disturbances. Ecological Indicators.DOI
Wenjing Chen, Huakun Zhou, Yang Wu, Jie Wang, Sha Xue (2026). Dominant role of abundant taxa in microbial community assembly regulating ecosystem function under plant functional group loss. Journal of Soils and Sediments.DOI
Xin Wang, Yan-Bin Wen, Hui-Yu Song, Nan Wang, Huakun Zhou, Gui-Ling Wu, Yan-Gong Du (2026). Precipitation and plant cover promote soil organic carbon accumulation in the northeastern Qinghai-Tibet Plateau. AoB Plants.DOI
Ming-Hua Song, Yuqiang Tian, Qi-Wu Hu, Yi-Kang Li, Xingliang Xu, Li-Li Zheng, J. H. C. Cornelissen, Huakun Zhou, Guiqiang Wang, Michel Loreau (2026). Nitrogen addition reduces alpine grassland stability via enhanced temporal synchrony and population variability. Oecologia.DOI
Qi-Nen He, Yingcheng Wang, Zhen Wang, Hui-Fang Shi, Guangxin Lu, Huakun Zhou (2026). Alpine meadow degradation reshapes diazotroph community assembly and enhances putative recruitment of diazotrophs from legume soils to sedge soils. Frontiers in Microbiology.DOI
An-Quan Xia, Tong Li, Huakun Zhou, T. Fu, Aizhen Liang, R. Pandey, Y. Dang (2026). Stop “Vandalising” Earth to Assist the Planet “Thrive”: Attaining Carbon Neutrality by Countering Land Degradation. Land Degradation & Development.DOI
Yanhe Wang, Mingxin Yang, Keyu Chen, Dongzhi Huangqing, Shouxing Wang, Xiaochun Ning, Hongkui Yang, Qiang Gu, Huakun Zhou (2025). Artificial perches improve the stability of soil microbial networks in degraded alpine steppe of Sanjiangyuan National Park. Journal Of Environmental Management, Vol. 395.DOI
Yingcheng Wang, Junbang Wang, Chu Wu, Huakun Zhou, Miao Zhang, Bo Zhao, Xi Peng, Xingsheng Yang, Xin Jin, Zhuohang Zhang, Guangxin Lu, Ye Deng (2025). Selective recruitment of core rhizobacterial communities with distinct life-history strategies by two grass genera. Journal Of Plant Ecology, Vol. 18, No. 6.DOI
Fawei Zhang, Hongqin Li, Jingbin Zhu, Chunyu Wang, Yunlong He, Juntao Zhu, Qiang Yu, Huakun Zhou, Yingnian Li, Naishen Liang (2025). Context dependencies in the responses of plant biomass and surface soil organic carbon content to nitrogen addition and precipitation change within alpine grasslands. Agriculture Ecosystems & Environment, Vol. 381.DOI
Xiaoyuan Yang, Huakun Zhou, Fanglin Liu (2025). Utilizing a vegetation restoration potential model to derive a reference for assessing ecological restoration of the Qinghai-Tibet Plateau. Ecological Engineering, Vol. 212.DOI
Fawei Zhang, Hongqin Li, Jingbin Zhu, Mengke Si, Bo Fan, Huakun Zhou, Yingnian Li (2025). Precipitation Determines the Spatial Variability of Vegetation and Topsoil Organic Carbon Densities of Alpine Grasslands in the Qinghai-Tibetan Plateau, China. Ecosystems, Vol. 280, No. 10.DOI
Mingyang Zhang, Yu Sun, Yong-Sheng Yang, Li Zhang, Zhenhua Zhang, Jing Zhao, Longfei Chen, Huakun Zhou (2025). Effects of alpine meadow degradation on soil nematode diversity and community composition on the Qinghai-Tibet Plateau. Ecological Indicators.DOI
Longxin Guo, Mingyang Zhang, Yong-Sheng Yang, Li Zhang, Yu Sun, Longfei Chen, Zijian Xu, Xiujuan Zhang, Zhenhua Zhang, Huakun Zhou, Haiming Ji (2025). Grazing regulates soil water-holding functions via altering plant functional groups in the southern Qilian Mountains. Science of the Total Environment.DOI
Yang Wu, HuaKun Zhou, WenJing Chen, HaoXiang Xue, HongFei Liu, Jie Wang, ShaoJuan Mao, GuoBin Liu, Sha Xue (2024). The combined nitrogen and phosphorus fertilizer application reduced soil multifunctionality in Qinghai-Tibet plateau grasslands, China. European Journal Of Soil Biology, Vol. 123.DOI
Mengyan Wang, Zuoyi Wang, Yuan Yang, Xinquan Zhao, Huakun Zhou, Shurong Zhou, Xin Yin, Yanmei Ren, Huanhuan Dong, Longxin Zhang, Zhen Ma, Chunhui Zhang (2024). Influences of elevational gradient on flower size and number of Gentiana lawrencei var. farreri. Ecology And Evolution, Vol. 14, No. 50.DOI
Jingjing Wei, Huakun Zhou, Xinqing Shao, Jian Sun, Li Ma, Zhonghua Zhang, Ruimin Qin, Hongye Su, Xue Hu, Tao Chang, Zhengchen Shi, Haze Ade, Huichun Wang (2024). Effects of short- and long-term plant functional group loss on alpine meadow community structure and soil nutrients. Ecology And Evolution, Vol. 14, No. 3.DOI
Guoxi Shi, Zhonghua Zhang, Li Ma, Yongjun Liu, Yibo Wang, Jean Yves Uwamungu, Huyuan Feng, Shikui Dong, Buqing Yao, Huakun Zhou (2024). Nitrogen addition drives changes in arbuscular mycorrhizal fungal richness through changes in plant species richness in revegetated alpine grassland. Fungal Ecology, Vol. 67.DOI
Gang Fu, Guang-Yu Zhang, Huakun Zhou (2024). Effects of long-term experimental warming on phyllosphere epiphytic bacterial and fungal communities of four alpine plants. Journal of Integrative Agriculture.DOI
Lei-Lei Qiao, Huakun Zhou, Zhanhui Wang, Wenjing Chen, Yuan-Ze Li, Yang Wu, Guo-bin Liu, Sha Xue (2024). Nine-year fertilization promoted C sink and mediated microbial nutrient utilization in alpine meadow soil aggregates. Agriculture, Ecosystems & Environment. Applied Soil Ecology.DOI
Yu Sun, Xiujuan Zhang, Yong-Sheng Yang, Yu-Heng Zhang, Junbang Wang, Mingyang Zhang, Chu Wu, Junliang Zou, Huakun Zhou, Jie-xia Li (2024). Alpine meadow degradation regulates soil microbial diversity via decreasing plant production on the Qinghai-Tibetan Plateau. Ecological Indicators.DOI
Yuan-Ze Li, Huakun Zhou, Qi-Fan Zhao, Wenjing Chen, Hong-Fei Liu, Bing Yang, Yang Wu, Guo-bin Liu, Sha Xue (2024). The Response of Grassland Production to N and P Fertilizer Management Mediated soil Nematode Community in Qinghai-Tibet Plateau of China. Journal of soil science and plant nutrition.DOI
Wenjing Chen, Huakun Zhou, Sha Xue (2024). Impact of Transition from Natural Grasslands Steppe to Monoculture Artificial Grasslands on Soil Food Webs in the Qinghai–Tibet Plateau. Agronomy.DOI
Yun Zhang, Yong-Sheng Yang, Bruce Osborne, Huakun Zhou, Juying Wu, Weiwei Zhang, Junliang Zou (2024). Edaphic factors control microbial biomass and elemental stoichiometry in alpine meadow soils of the Tibet Plateau. Plant and Soil.DOI
Zi-Wen Zhao, Huakun Zhou, Wenjing Chen, Yang Wu, Guo-bin Liu, Sha Xue (2024). Response of soil structure and fertility to long-term fertilization in alpine grasslands revealed based on fractal theory. Soil & Tillage Research.DOI
Yandi She, Xi-Lai Li, Jing Zhang, Huakun Zhou (2024). Effects of soil characteristics on grassland productivity in long-term artificial grassland establishment. Global Ecology and Conservation.DOI
Wen-geng Qin, Jiguang Feng, Qiufang Zhang, Xiao-Cun Yuan, Fei Ren, Huakun Zhou, Biao Zhu (2024). Different responses of priming effects in long-term nitrogen- and phosphorus-fertilized soils to exogenous carbon inputs. Plant and Soil.DOI
Ming-Hua Song, Yu-Qiang Tian, Yi-Kang Li, Xingliang Xu, Li-Li Zheng, Huakun Zhou, Y. Kuzyakov, Guiqiang Wang, Yang-Jian Zhang, J. H. C. Cornelissen (2024). Loss of taxonomic and functional diversity and decrease in primary productivity with nitrogen enrichment after a long-term release from grazing in an alpine grassland. Science of the Total Environment.DOI
Wenjing Chen, Huakun Zhou, Yang Wu, Jie Wang, Ziwen Zhao, Yuanze Li, Leilei Qiao, Kelu Chen, Guobin Liu, Coen Ritsema, Violette Geissen, Xue Sha (2023). Effects of deterministic assembly of communities caused by global warming on coexistence patterns and ecosystem functions. Journal Of Environmental Management, Vol. 345.DOI
Wenjin Li, Johannes M. H. Knops, Xi Zhou, Huijun Jin, Zhiqiang Xiang, Cairang Ka Zhuo, Xiaoying Jin, Huakun Zhou, ShiKui Dong (2023). [Review] Anchoring grassland sustainability with a nature-based small burrowing mammal control strategy. Journal Of Animal Ecology, Vol. 920, No. 70.DOI
Wenlong Li, Xuejing Shang, Hepiao Yan, Jing Xu, Tiangang Liang, Huakun Zhou (2023). Impact of restoration measures on plant and soil characteristics in the degraded alpine grasslands of the Qinghai Tibetan Plateau: A meta-analysis. Agriculture Ecosystems & Environment, Vol. 347.DOI
Hongye Su, Li Ma, Tao Chang, Ruimin Qin, Zhonghua Zhang, Yandi She, Jingjing Wei, Chenyu Zhou, Xue Hu, Zhengchen Shi, Haze Adi, Honglin Li, Huakun Zhou (2023). Effects of Main Land-Use Types on Plant and Microbial Diversity and Ecosystem Multifunctionality in Degraded Alpine Grasslands. Land, Vol. 12, No. 3.DOI
Wen-geng Qin, Jiguang Feng, Qiufang Zhang, Xiao-Cun Yuan, Huakun Zhou, Biao Zhu (2023). Nitrogen and phosphorus addition mediate soil priming effects via affecting microbial stoichiometric balance in an alpine meadow. Science of the Total Environment.DOI
Yuanze Li, Huakun Zhou, Wenjing Chen, Yang Wu, LeiLei Qiao, ZiRan Yan, GuoBin Liu, Sha Xue (2022). Long-term warming does not affect soil ecoenzyme activity and original microbial nutrient limitation on the Qinghai-Tibet Plateau. Soil Ecology Letters, Vol. 40, No. 40.DOI
Lina Shi, Zhenrong Lin, Shiming Tang, Cuoji Peng, Zeying Yao, Qing Xiao, Huakun Zhou, Kesi Liu, Xinqing Shao (2022). Interactive effects of warming and managements on carbon fluxes in grasslands: A global meta-analysis. Agriculture Ecosystems & Environment, Vol. 340.DOI
Lei Wang, Xufeng Mao, Xiuhua Song, Wenjia Tang, Wenying Wang, Hongyan Yu, Yanfang Deng, Ziping Zhang, Zhijun Zhang, Huakun Zhou (2022). How Rising Water Levels Altered Ecosystem Provisioning Services of the Area around Qinghai Lake from 2000 to 2020: An InVEST-RF-GTWR Combined Method. Land, Vol. 11, No. 9.DOI
Shan Luo, Richard D. Bardgett, Bernhard Schmid, David Johnson, Kenny Png, Urs Schaffner, Huakun Zhou, Buqing Yao, Xiangyang Hou, Nicholas J. Ostle (2022). Historical context modifies plant diversity-community productivity relationships in alpine grassland. Journal Of Ecology, Vol. 1100, No. 9.DOI
Huidan He, Jingbin Zhu, Yangong Du, Jiapeng Qu, Chen Kelong, Huakun Zhou (2022). Effects of Heavy Degradation on Alpine Meadows: Soil N2O Emission Rates and Meta-Analysis in the Tibetan Plateau. Land, Vol. 11, No. 8.DOI
Lingling Tong, Xufeng Mao, Xiuhua Song, Xiaoyan Wei, Wenjia Tang, Yanfang Deng, Hongyan Yu, Zhuo Deng, Feng Xiao, Huakun Zhou, Xinan Yin (2022). PSR-BP Neural Network-Based Health Assessment of the Huangshui Plateau Urban Wetlands in China. Frontiers In Ecology And Evolution, Vol. 10.DOI
Hongqin Li, Jingbin Zhu, Fawei Zhang, Gang Qin, Yongsheng Yang, Yikang Li, Junbang Wang, Guangmin Cao, Yingnian Li, Huakun Zhou, Mingyuan Du (2022). The Predominance of Nongrowing Season Emissions to the Annual Methane Budget of a Semiarid Alpine Meadow on the Northeastern Qinghai-Tibetan Plateau. Ecosystems, Vol. 250, No. 30.DOI
Ying Chen, Mengguang Han, Xia Yuan, Yanhui Hou, Wenkuan Qin, Huakun Zhou, Xinquan Zhao, Julia A. Klein, Biao Zhu (2022). Warming has a minor effect on surface soil organic carbon in alpine meadow ecosystems on the Qinghai-Tibetan Plateau. Global Change Biology, Vol. 280, No. 40.DOI
Ziqi Guo, Huakun Zhou, Wenjing Chen, Yang Wu, Yuan-Ze Li, Lei-Lei Qiao, Qi-Ming You, Guo-bin Liu, Sha Xue (2022). Impacts of 21-year field warming on soil erodibility in the Qinghai-Tibetan Plateau, China. Geoderma.DOI
Yunying Wang, Weiwei Pei, Guangmin Cao, Xiaowei Guo, Huakun Zhou, Yangong Du (2021). Moderate Grazing Increases Water Use Efficiency for Environmental Health in Alpine Meadows of the Tibetan Plateau. Frontiers In Ecology And Evolution, Vol. 9.DOI
Zhe Chen, Shidong Ge, Zhenhua Zhang, Yangong Du, Buqing Yao, Huichun Xie, Pan Liu, Yufang Zhang, Wenying Wang, Huakun Zhou (2021). Soil Moisture but Not Warming Dominates Nitrous Oxide Emissions During Freeze-Thaw Cycles in a Qinghai-Tibetan Plateau Alpine Meadow With Discontinuous Permafrost. Frontiers In Ecology And Evolution, Vol. 9.DOI
Hongqin Li, Fawei Zhang, Jingbin Zhu, Xiaowei Guo, Yikang Li, Li Lin, Leiming Zhang, Yongsheng Yang, Yingnian Li, Guangmin Cao, Huakun Zhou, Mingyuan Du (2021). Precipitation rather than evapotranspiration determines the warm-season water supply in an alpine shrub and an alpine meadow. Agricultural And Forest Meteorology, Vol. 300.DOI
Jiapeng Qu, Zelin Liu, Zhenggang Guo, Yikang Li, Huakun Zhou (2021). A System Dynamics Model for Assessing the Efficacy of Lethal Control for Sustainable Management of Ochotona curzoniae on Tibetan Plateau. Sustainability, Vol. 13, No. 2.DOI
Xiaoting Wei, Yanan Shi, Fuwen Qin, Huakun Zhou, Xinqing Shao (2021). Effects of experimental warming, precipitation increase and their interaction on AM fungal community in an alpine grassland of the Qinghai-Tibetan Plateau. European Journal Of Soil Biology, Vol. 102.DOI
Yudan Xu, Shikui Dong, Xiaoxia Gao, Mingyue Yang, Shuai Li, Hao Shen, Jiannan Xiao, Yuhui Han, Jing Zhang, Yu Li, Yangliu Zhi, Yunfeng Yang, Shiliang Liu, Quanming Dong, Huakun Zhou (2021). Aboveground community composition and soil moisture play determining roles in restoring ecosystem multifunctionality of alpine steppe on Qinghai-Tibetan Plateau. Agriculture Ecosystems & Environment, Vol. 305.DOI
Pan Liu, Wen-Ying Wang, Huakun Zhou, Chong Yang (2021). Ammonia volatilisation and N recovery of nitrogen fertiliser on former alpine grassland based on a nitrogen isotope labelling technique.DOI
Dangjun Wang, Huakun Zhou, Buqing Yao, Wenying Wang, Shikui Dong, Zhanhuan Shang, Yandi She, Li Ma, Xiaotao Huang, Zhonghua Zhang, Qian Zhang, Fengyu Zhao, Juan Zuo, Zhun Mao (2020). Effects of nutrient addition on degraded alpine grasslands of the Qinghai-Tibetan Plateau: A meta-analysis. Agriculture Ecosystems & Environment, Vol. 301.DOI
Zhenchao Zhang, Jian Sun, Miao Liu, Ming Xu, Yi Wang, Gao-lin Wu, Huakun Zhou, Chongchong Ye, Dorji Tsechoe, Tianxing Wei (2020). [Review] Don't judge toxic weeds on whether they are native but on their ecological effects. Ecology And Evolution, Vol. 100, No. 17.DOI
Yangong Du, Xun Ke, Licong Dai, Guangmin Cao, Huakun Zhou, Xiaowei Guo (2020). Moderate grazing increased alpine meadow soils bacterial abundance and diversity index on the Tibetan Plateau. Ecology And Evolution, Vol. 10, No. 16.DOI
Jing Guo, Jun Ren, Xiaotao Huang, Guifang He, Yan Shi, Huakun Zhou (2020). The Dynamic Evolution of the Ecological Footprint and Ecological Capacity of Qinghai Province. Sustainability, Vol. 12, No. 7.DOI
Tiancai Zhou, Miao Liu, Jian Sun, Yurui Li, Peili Shi, Atsushi Tsunekawa, Huakun Zhou, Shuhua Yi, Xian Xue (2020). The patterns and mechanisms of precipitation use efficiency in alpine grasslands on the Tibetan Plateau. Agriculture Ecosystems & Environment, Vol. 292.DOI
Xiaowei Guo, Licong Dai, Qian Li, Dawen Qian, Guangmin Cao, Huakun Zhou, Yangong Du (2020). Light Grazing Significantly Reduces Soil Water Storage in Alpine Grasslands on the Qinghai-Tibet Plateau. Sustainability, Vol. 120, No. 60.DOI
Lv Wangwang, Luo Caiyun, Zhang Lirong, Niu Haishan, Zhang Zhenhua, Wang Shiping, Wang Yanfen, Jiang Lili, Wang Yonghui, He Jinsheng, Kardol Paul, Wang Qi, Li Bowen, Liu Peipei, Dorji Tsechoe, Zhou Huakun, Zhao Xinquan, Zhao Liang (2020). Net neutral carbon responses to warming and grazing in alpine grassland ecosystems. Agricultural And Forest Meteorology, Vol. 280.DOI
Licong Dai, Xiaowei Guo, Xun Ke, Yuting Lan, Fawei Zhang, Yikang Li, Li Lin, Qian Li, Guangmin Cao, Bo Fan, Dawen Qian, Huakun Zhou, Yangong Du (2020). Biomass allocation and productivity-richness relationship across four grassland types at the Qinghai Plateau. Ecology And Evolution, Vol. 10, No. 1.DOI
Qiaoyi Hua, Yi Yu, Shikui Dong, Shuai Li, Hao Shen, Yuhui Han, Jing Zhang, Jianan Xiao, Shiliang Liu, Quanming Dong, Huakun Zhou, Kelly Wessell (2019). Leaf spectral responses of Poa crymophila to nitrogen deposition and climate change on Qinghai-Tibetan Plateau. Agriculture Ecosystems & Environment, Vol. 284.DOI
Bingrong Zhou, Shuai Li, Fu Li, Shikui Dong, Fulin Ma, Shengcui Zhu, Huakun Zhou, Paul Stufkens (2019). Plant functional groups asynchrony keep the community biomass stability along with the climate change- a 20-year experimental observation of alpine meadow in eastern Qinghai-Tibet Plateau. Agriculture Ecosystems & Environment, Vol. 282.DOI
Zhen Ma, Charles G. Willis, Chunhui Zhang, Huakun Zhou, Xinquan Zhao, Shikui Dong, Buqing Yao, Xiaotao Huang, Feng-Yu Zhao, Guang-Jin Yin, Dengxian Wei, Guozhen Du (2019). Direct and indirect effect of seed size on seedling survival along an experimental light availability gradient. Agriculture Ecosystems & Environment, Vol. 281.DOI
Xiaoxia Gao, Shikui Dong, Yudan Xu, Shengnan Wu, Xiaohui Wu, Xi Zhang, Yangliu Zhi, Shuai Li, Shiliang Liu, Yu Li, Zhanhuan Shang, Quanmin Dong, Huakun Zhou, Paul Stufkens (2019). Resilience of revegetated grassland for restoring severely degraded alpine meadows is driven by plant and soil quality along recovery time: A case study from the Three-river Headwater Area of Qinghai-Tibetan Plateau. Agriculture Ecosystems & Environment, Vol. 279.DOI
Xin Li, Xinran Ke, Huakun Zhou, Yanhong Tang (2019). Contrasting altitudinal patterns of leaf UV reflectance and absorbance in four herbaceous species on the Qinghai-Tibetan Plateau. Journal Of Plant Ecology, Vol. 12, No. 2.DOI
Fangping Wang, Guoxi Shi, Ostle Nicholas, Buqing Yao, Mingfei Ji, Wenying Wang, Zhen Ma, Huakun Zhou, Xinquan Zhao (2018). Ecosystem nitrogen retention is regulated by plant community trait interactions with nutrient status in an alpine meadow. Journal Of Ecology, Vol. 106, No. 4.DOI
Fei Ren, Weimin Song, Litong Chen, Zhaorong Mi, Zhenhua Zhang, Wenyan Zhu, Huakun Zhou, Guangmin Cao, Jin-Sheng He (2017). Phosphorus does not alleviate the negative effect of nitrogen enrichment on legume performance in an alpine grassland. Journal Of Plant Ecology, Vol. 10, No. 5.DOI
Chunhui Zhang, Charles G. Willis, Julia A. Klein, Zhen Ma, Junyong Li, Huakun Zhou, Xinquan Zhao (2017). Recovery of plant species diversity during long-term experimental warming of a species-rich alpine meadow community on the Qinghai-Tibet plateau. Biological Conservation, Vol. 213.DOI
Yushan Long, Zhou Li, Wei Liu, Huakun Zhou (2010). Using Remote Sensing and GIS Technologies to Estimate Grass Yield and Livestock Carrying Capacity of Alpine Grasslands in Golog Prefecture, China.DOI
Liang Zhao, Yingnian Li, Shixiao Xu, Huakun Zhou, Song Gu, Guirui Yu, Xinquan Zhao (2006). Diurnal, seasonal and annual variation in net ecosystem CO2 exchange of an alpine shrubland on Qinghai-Tibetan plateau. Global Change Biology, Vol. 120, No. 100.DOI
Li-Cong Dai, Ruiyu Fu, Xiao-Wei Guo, Yangong Du, G. Cao, Huakun Zhou, Zhong-Min Hu. Biocrust reduced soil water retention and soil infiltration in the alpine Kobresia meadow.